December 30 , 2016:

Pterosaurs tracks in the Cameros basin are plentiful and assorted. This fact has allowed to define several Pteraichnus ichnospecies and moreover to distinguish other morphotypes. The study of the new tracksite of La Muela (Soria, Spain) describes Pteraichnus cf. stokesi ichnites that is an unknown ichnospecies until now and that confirms the wide diversity of this type of tracks in the Cameros Basin. Their characteristics correspond to the ones of the Upper Jurassic tracksites of United States. Similar tracks have already been described in other tracksites, both inside and outside the Iberian Peninsula during the Upper Jurassic-Lower Cretaceous transit. Because of their shape and morphometrical characteristics they can be related to the pterosaurs of the Archaeopterodactyloidea clade. The analysis of this ichnogenus indicates the need for a deep review because encompasses ichnites with a big variety of shapes and morphometric characteristics.

Carlos Pascual-Arribas and Nieves Hernández-Medrano (2016)
Huellas de Pteraichnus en La Muela (Soria, España): consideraciones sobre el icnogénero y sobre la diversidad de huellas de pterosaurios en la Cuenca de Cameros.
[Pteraichnus tracks in La Muela (Soria, Spain): considerations on the ichnogenus and diversity of pterosaur tracks in the Cameros Basin.]
Revista de la Sociedad Geológica de España 29 (2): 89-105
ISSN (versión impresa): 0214-2708
ISSN (Internet): 2255-1379